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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
No data
RNA expressioni
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression levels for human tissues was created by combining the data from HPA and GTEx datasets.
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Consensus:
67.2
nTPM
HPA Brain:
29.2
nTPM
GTEx:
67.2
nTPM
FANTOM5:
17.8
Scaled Tags Per Million
Pig:
33.5
nTPM
Mouse:
11.9
nTPM
HIPPOCAMPAL FORMATION - Annotated protein expression
Pending normal tissue analysis
HIPPOCAMPAL FORMATION - HPA Brain RNA-seq
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
29.2
Sample id
Sample description
nTPM
Hippocampus nTPM: 26.0
2621
Female, age 67, right hemisphere
29.2
2529
Female, age 94, left hemisphere
28.9
1898
Male, age 80, left hemisphere
27.2
1855
Male, age 76, left hemisphere
26.4
2148
Female, age 91, left hemisphere
23.7
1911
Female, age 85, right hemisphere
20.7
Dentate gyrus nTPM: 21.9
2010
Male, age 76, both hemispheres
23.3
2530
Female, age 94, left hemisphere
23.3
2246
Female, age 85, both hemispheres
22.1
2531
Female, age 67, both hemispheres
21.9
1927
Female, age 91, both hemispheres
21.4
2528
Male, age 74, both hemispheres
19.6
Entorhinal gyrus nTPM: 28.7
1938
Female, age 85, left hemisphere
31.2
1787
Male, age 76, left hemisphere
30.3
2764
Male, age 80, right hemisphere
30.2
2613
Female, age 94, left hemisphere
29.4
2686
Male, age 75, right hemisphere
27.7
2117
Female, age 91, right hemisphere
23.6
Hippocampus, CA1 nTPM: 23.5
2749
Male, age 80, left hemisphere
26.2
2082
Female, age 91, right hemisphere
25.5
2527
Male, age 74, both hemispheres
23.2
2045
Male, age 76, both hemispheres
23.1
1873
Female, age 85, both hemispheres
19.4
Hippocampus, CA2 nTPM: 24.2
1940
Female, age 85, left hemisphere
26.4
1892
Male, age 76, both hemispheres
25.4
2113
Female, age 91, right hemisphere
25.0
2437
Female, age 94, left hemisphere
24.1
2249
Male, age 74, both hemispheres
23.1
1963
Male, age 80, left hemisphere
21.2
Hippocampus, CA3 nTPM: 24.3
2137
Male, age 80, left hemisphere
26.8
1867
Male, age 76, both hemispheres
25.4
2861
Female, age 67, both hemispheres
24.6
1891
Female, age 85, both hemispheres
24.2
2436
Male, age 74, both hemispheres
22.8
2620
Female, age 94, left hemisphere
22.2
Parahippocampal cortex nTPM: 27.0
1850
Male, age 80, both hemispheres
33.5
2245
Female, age 91, both hemispheres
30.1
2622
Female, age 67, right hemisphere
29.0
1990
Female, age 91, right hemisphere
27.6
2692
Male, age 74, both hemispheres
27.4
2532
Female, age 72, left hemisphere
27.3
2693
Female, age 94, both hemispheres
25.8
2250
Female, age 67, right hemisphere
25.7
2018
Male, age 76, both hemispheres
23.0
1933
Female, age 85, both hemispheres
20.9
Perirhinal gyrus nTPM: 29.2
1847
Female, age 91, left hemisphere
32.6
1794
Male, age 80, right hemisphere
29.8
1841
Female, age 85, right hemisphere
29.0
2848
Male, age 76, right hemisphere
28.1
2521
Female, age 94, right hemisphere
26.5
Subiculum nTPM: 26.9
2438
Female, age 94, left hemisphere
34.8
1977
Female, age 85, both hemispheres
27.7
2619
Male, age 74, both hemispheres
26.0
2042
Male, age 76, both hemispheres
25.7
1901
Male, age 80, both hemispheres
25.4
2729
Female, age 91, right hemisphere
22.1
HIPPOCAMPAL FORMATION - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as mean nTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
67.2
GTEx sample id
Sample description
nTPM
Brain - Hippocampus Average nTPM: 67.2
Samples: 197
Max nTPM: 164.0
Min nTPM: 35.1
Std nTPM: 18.0
Median nTPM: 66.8
GTEX-X4EP-0011-R1A-SM-3P624
60-69 years, female
164.0
GTEX-1HR98-0011-R1a-SM-ARZLQ
60-69 years, male
127.4
GTEX-18A6Q-0011-R1b-SM-731DI
60-69 years, male
121.8
GTEX-ZF28-0011-R1a-SM-4WWD1
60-69 years, male
113.2
GTEX-16NPV-0011-R1b-SM-7MGVZ
50-59 years, female
111.4
GTEX-NL3H-0011-R1a-SM-48TDJ
60-69 years, male
110.4
GTEX-13PDP-0011-R1a-SM-5PNX5
60-69 years, male
103.1
GTEX-1EWIQ-0011-R1a-SM-9OSX7
70-79 years, female
102.9
GTEX-ZE7O-0011-R1a-SM-57WDM
50-59 years, female
100.9
GTEX-1H4P4-0011-R1b-SM-CL54V
50-59 years, male
99.9
GTEX-OHPN-0011-R1A-SM-2I5GB
60-69 years, female
98.9
GTEX-13FTY-0011-R1a-SM-5LZXE
40-49 years, female
95.3
GTEX-13FHP-0011-R1b-SM-5K7XL
50-59 years, male
94.5
GTEX-1C6VQ-0011-R1a-SM-7P8Q2
50-59 years, male
93.2
GTEX-13NYS-0011-R1b-SM-5MR55
60-69 years, male
93.0
GTEX-1HSMO-0011-R1a-SM-CM2SV
60-69 years, male
92.7
GTEX-17HII-0011-R1b-SM-7MGVW
50-59 years, male
92.1
GTEX-139T8-0011-R1a-SM-5HL75
60-69 years, male
91.4
GTEX-17F97-0011-R1b-SM-7IGM9
50-59 years, male
90.7
GTEX-183WM-0011-R1a-SM-718A8
70-79 years, female
89.6
GTEX-1H3VE-0011-R1a-SM-A9G1K
40-49 years, male
88.7
GTEX-1GN2E-0011-R1b-SM-CE6RJ
60-69 years, male
88.6
GTEX-1H23P-0011-R1a-SM-ACKVC
60-69 years, male
88.5
GTEX-QMR6-0011-R1A-SM-32PKW
50-59 years, male
88.2
GTEX-13OW5-0011-R1b-SM-5L3GE
60-69 years, male
87.1
GTEX-14BIM-0011-R1a-SM-69LQI
60-69 years, female
84.9
GTEX-1B996-0011-R1b-SM-7P8PO
60-69 years, male
84.0
GTEX-1CB4J-0011-R1a-SM-9WG64
60-69 years, male
83.9
GTEX-17HHY-0011-R1a-SM-7189V
50-59 years, male
83.6
GTEX-1H1DG-0011-R1b-SM-CKZP1
70-79 years, male
83.4
GTEX-1J8Q2-0011-R1a-SM-D4P2D
60-69 years, male
83.4
GTEX-1C64N-0011-R1b-SM-7P8Q7
50-59 years, male
82.4
GTEX-11UD1-0011-R1b-SM-5BC7D
40-49 years, female
82.3
GTEX-16GPK-0011-R1b-SM-7EWDH
60-69 years, male
81.3
GTEX-11EI6-0011-R1a-SM-DO93L
60-69 years, male
80.1
GTEX-XMD1-0011-R1A-SM-4AT4C
60-69 years, male
79.5
GTEX-1IOXB-0011-R1a-SM-D4P27
50-59 years, male
79.5
GTEX-TSE9-0011-R1A-SM-3DB7E
60-69 years, female
79.4
GTEX-1A32A-0011-R1a-SM-7P8SQ
50-59 years, female
79.3
GTEX-1C6VR-0011-R1a-SM-7MXUE
60-69 years, male
79.3
GTEX-13FHO-0011-R1b-SM-5KLZG
60-69 years, male
78.9
GTEX-13QJC-0011-R1b-SM-5S2UU
60-69 years, female
77.9
GTEX-11ZTS-0011-R1a-SM-5BC71
60-69 years, female
77.1
GTEX-1GF9V-0011-R1a-SM-CKZNQ
60-69 years, male
77.0
GTEX-13RTL-0011-R1a-SM-5PNZ8
40-49 years, male
76.9
GTEX-1HGF4-0011-R1a-SM-CJI3S
60-69 years, male
76.8
GTEX-1EKGG-0011-R1b-SM-9QEI1
60-69 years, male
76.7
GTEX-1GF9W-0011-R1a-SM-9QEIQ
60-69 years, male
76.5
GTEX-1I1GS-0011-R1a-SM-CJI43
60-69 years, male
76.0
GTEX-PWO3-0011-R1A-SM-2I5EW
60-69 years, female
75.7
GTEX-15UF6-0011-R1a-SM-9QEGH
70-79 years, female
75.4
GTEX-13NZ8-0011-R1b-SM-5KM3V
60-69 years, female
75.0
GTEX-14LLW-0011-R1a-SM-5ZZVD
70-79 years, female
74.5
GTEX-1EX96-0011-R1a-SM-CE6R5
50-59 years, male
74.5
GTEX-17MF6-0011-R1a-SM-72D6O
40-49 years, male
74.3
GTEX-11TUW-0011-R1a-SM-DO92S
60-69 years, male
74.2
GTEX-1GTWX-0011-R1a-SM-CE6RL
60-69 years, male
74.2
GTEX-ZVZQ-0011-R1a-SM-57WBU
60-69 years, female
73.9
GTEX-13NYB-0011-R1b-SM-5KM4F
40-49 years, male
73.7
GTEX-15CHQ-0011-R1a-SM-COH27
60-69 years, male
73.7
GTEX-13QIC-0011-R1a-SM-5O9CJ
60-69 years, female
73.2
GTEX-144GL-0011-R1a-SM-5PNZ5
50-59 years, male
73.1
GTEX-1399T-0011-R1b-SM-5DUWN
60-69 years, male
72.9
GTEX-Y8DK-0011-R1A-SM-4RGLO
60-69 years, male
72.8
GTEX-14PJM-0011-R1a-SM-CNPO3
50-59 years, female
72.5
GTEX-1B8SF-0011-R1b-SM-7P8Q8
60-69 years, male
72.5
GTEX-1CAV2-0011-R1a-SM-ARZML
40-49 years, female
72.2
GTEX-1J1OQ-0011-R1a-SM-A9G1Z
40-49 years, female
72.2
GTEX-132Q8-0011-R1a-SM-5EGLK
60-69 years, male
72.0
GTEX-NPJ7-0011-R1a-SM-3GACT
60-69 years, female
71.2
GTEX-11DXW-0011-R1a-SM-DNZZD
40-49 years, male
71.0
GTEX-13O3Q-0011-R1b-SM-5KM3L
50-59 years, male
70.5
GTEX-1LG7Y-0011-R1a-SM-EYYWV
50-59 years, male
70.3
GTEX-12WSD-0011-R1b-SM-5LZVY
60-69 years, female
69.8
GTEX-16Z82-0011-R1b-SM-7EPHZ
60-69 years, male
69.5
GTEX-144GO-0011-R1a-SM-5PNWO
60-69 years, male
69.3
GTEX-1E1VI-0011-R1a-SM-ARZMM
50-59 years, male
69.2
GTEX-1I1GV-0011-R1a-SM-CL559
60-69 years, male
69.1
GTEX-11WQC-0011-R1b-SM-DO932
60-69 years, male
68.8
GTEX-1EH9U-0011-R1b-SM-CJI2B
60-69 years, male
68.4
GTEX-14PQA-0011-R1b-SM-6EU1G
50-59 years, female
68.2
GTEX-P44G-0011-R1A-SM-2I3FE
30-39 years, male
68.2
GTEX-13JVG-0011-R1a-SM-5KM32
60-69 years, male
67.9
GTEX-1A8G6-0011-R1a-SM-7P8PC
60-69 years, male
67.9
GTEX-12WSF-0011-R1a-SM-5DUVI
70-79 years, male
67.7
GTEX-X4XX-0011-R1B-SM-3P622
60-69 years, male
67.7
GTEX-1F6I4-0011-R1a-SM-9OSWU
60-69 years, male
67.7
GTEX-13N1W-0011-R1b-SM-5MR4T
70-79 years, male
67.6
GTEX-17HG3-0011-R1a-SM-7LG6H
30-39 years, female
67.6
GTEX-1E2YA-0011-R1b-SM-CJI29
50-59 years, male
67.6
GTEX-1H3NZ-0011-R1a-SM-CKZP5
60-69 years, male
67.5
GTEX-13CF2-0011-R1a-SM-5LZY1
60-69 years, male
67.4
GTEX-13FXS-0011-R1a-SM-5K7U6
50-59 years, male
67.4
GTEX-14LZ3-0011-R1a-SM-69LP5
60-69 years, female
67.4
GTEX-1HBPI-0011-R1b-SM-A9SKF
50-59 years, male
67.3
GTEX-13OW8-0011-R1a-SM-5L3HI
60-69 years, male
67.2
GTEX-1H1CY-0011-R1a-SM-CKZOZ
60-69 years, female
67.1
GTEX-1F75W-0011-R1a-SM-ARU81
60-69 years, male
67.0
GTEX-1HT8W-0011-R1b-SM-CJI3V
30-39 years, male
66.8
GTEX-1H1ZS-0011-R1b-SM-A9G1E
70-79 years, male
66.3
GTEX-1GZ2Q-0011-R1a-SM-CJI2Y
40-49 years, male
66.0
GTEX-1GN1U-0011-R1a-SM-CJI25
50-59 years, female
65.8
GTEX-13S7M-0011-R1b-SM-5PNZE
60-69 years, female
65.2
GTEX-1GMR8-0011-R1b-SM-CJI2O
50-59 years, female
65.0
GTEX-1LB8K-0011-R1b-SM-EZ6MV
50-59 years, male
64.8
GTEX-13112-0011-R1b-SM-5HL7K
50-59 years, male
64.6
GTEX-WVLH-0011-R1A-SM-4MVOK
50-59 years, male
64.3
GTEX-12WSM-0011-R1a-SM-5LZW6
60-69 years, male
64.1
GTEX-145LU-0011-R1a-SM-5Q5CE
50-59 years, male
64.0
GTEX-13OW7-0011-R1b-SM-5L3HF
50-59 years, male
63.8
GTEX-1F48J-0011-R1a-SM-9QEIC
50-59 years, female
63.8
GTEX-UTHO-0011-R1A-SM-3GIJO
60-69 years, male
63.7
GTEX-13NZA-0011-R1b-SM-5LUAF
60-69 years, male
63.1
GTEX-13OVL-0011-R1a-SM-5L3H5
50-59 years, male
62.8
GTEX-12WSE-0011-R1b-SM-5GU4V
20-29 years, male
62.7
GTEX-N7MT-0011-R1a-SM-CNPO1
60-69 years, female
62.1
GTEX-12WSH-0011-R1a-SM-5GU5V
50-59 years, male
62.0
GTEX-1B933-0011-R1b-SM-9QEGW
50-59 years, female
61.1
GTEX-1GZHY-0011-R1a-SM-CE6RR
50-59 years, female
61.0
GTEX-13OW6-0011-R1a-SM-5L3HQ
50-59 years, male
60.7
GTEX-13SLW-0011-R1a-SM-5S2W7
70-79 years, male
60.7
GTEX-11GSP-0011-R1a-SM-9QEJ3
60-69 years, female
60.4
GTEX-17EVP-0011-R1a-SM-7DHLA
40-49 years, male
60.3
GTEX-1CB4G-0011-R1b-SM-9QEHT
60-69 years, male
60.3
GTEX-S7SE-0011-R1A-SM-2XCDE
50-59 years, male
60.0
GTEX-1I1HK-0011-R1b-SM-CM2T7
60-69 years, male
59.9
GTEX-1F75A-0011-R1b-SM-CE6RA
50-59 years, male
59.6
GTEX-1GN73-0011-R1a-SM-CKZNT
60-69 years, male
59.6
GTEX-1A3MX-0011-R1b-SM-7P8PH
60-69 years, male
59.3
GTEX-17EVQ-0011-R1a-SM-7189S
70-79 years, male
59.1
GTEX-11GS4-0011-R1a-SM-DO129
60-69 years, male
58.9
GTEX-14PN4-0011-R1b-SM-69LP8
50-59 years, male
58.4
GTEX-YJ89-0011-R1a-SM-4RGLS
60-69 years, male
58.0
GTEX-13RTJ-0011-R1a-SM-5O9D9
60-69 years, male
57.5
GTEX-ZAB4-0011-R1a-SM-4RGNO
40-49 years, male
57.4
GTEX-14E6D-0011-R1b-SM-5YYAW
50-59 years, male
57.3
GTEX-17JCI-0011-R1b-SM-718A3
60-69 years, female
57.0
GTEX-P44H-0011-R1A-SM-3NM8J
40-49 years, male
57.0
GTEX-T6MN-0011-R1A-SM-32QOY
50-59 years, male
56.9
GTEX-1BAJH-0011-R1a-SM-7P8SV
70-79 years, male
56.7
GTEX-13O3O-0011-R1b-SM-5KM2K
60-69 years, female
56.5
GTEX-14PJO-0011-R1b-SM-6AJAK
60-69 years, male
56.2
GTEX-WWYW-0011-R1A-SM-3TW8G
50-59 years, female
56.0
GTEX-1ICLZ-0011-R1b-SM-A9SM3
60-69 years, male
55.9
GTEX-11GSO-0011-R1b-SM-57WD3
60-69 years, male
55.1
GTEX-N7MS-0011-R1a-SM-2HMJG
60-69 years, male
54.8
GTEX-1I1GQ-0011-R1a-SM-CY8GG
50-59 years, male
54.6
GTEX-15ER7-0011-R1b-SM-7KUE1
20-29 years, female
54.5
GTEX-NPJ8-0011-R1a-SM-33HCB
40-49 years, male
53.6
GTEX-1AX9I-0011-R1b-SM-7RHG2
60-69 years, male
53.5
GTEX-14C5O-0011-R1b-SM-6AJBW
60-69 years, male
53.0
GTEX-139TT-0011-R1a-SM-5LZVD
60-69 years, male
52.9
GTEX-13OVJ-0011-R1b-SM-5L3I6
50-59 years, female
52.1
GTEX-1IKK5-0011-R1b-SM-CE6SQ
60-69 years, male
52.1
GTEX-13FLW-0011-R1b-SM-5LZX2
50-59 years, male
52.0
GTEX-145LS-0011-R1b-SM-5PNUP
60-69 years, female
51.8
GTEX-X585-0011-R1B-SM-46MVE
50-59 years, male
51.7
GTEX-14ABY-0011-R1a-SM-6EU2W
50-59 years, male
51.6
GTEX-QVUS-0011-R1A-SM-3GAD2
60-69 years, female
51.5
GTEX-1212Z-0011-R1b-SM-5BC72
60-69 years, male
50.9
GTEX-13N2G-0011-R1b-SM-5MR3G
60-69 years, male
50.7
GTEX-15DCD-0011-R1b-SM-6LPIJ
70-79 years, female
50.7
GTEX-14ASI-0011-R1b-SM-5S2TY
60-69 years, male
50.6
GTEX-WL46-0011-R1A-SM-3LK6M
50-59 years, male
50.6
GTEX-11ONC-0011-R1a-SM-57WD4
60-69 years, male
50.3
GTEX-ZDXO-0011-R1a-SM-4WKF4
60-69 years, male
50.1
GTEX-WHSE-0011-R1A-SM-3P5ZK
20-29 years, male
49.6
GTEX-1B8L1-0011-R1a-SM-7P8SS
60-69 years, male
49.2
GTEX-1H3O1-0011-R1b-SM-CL554
60-69 years, male
48.8
GTEX-QVJO-0011-R1A-SM-2S1QI
60-69 years, female
48.6
GTEX-WZTO-0011-R1B-SM-3NMAR
40-49 years, male
48.5
GTEX-1HSKV-0011-R1b-SM-CMKH7
60-69 years, male
48.4
GTEX-1313W-0011-R1b-SM-5EQ4A
50-59 years, female
48.3
GTEX-11ZVC-0011-R1b-SM-5BC6M
50-59 years, female
47.8
GTEX-16XZZ-0011-R1a-SM-7LTAH
60-69 years, male
47.4
GTEX-13X6I-0011-R1b-SM-5PNZC
60-69 years, male
47.2
GTEX-QDT8-0011-R1A-SM-32PKS
30-39 years, female
46.7
GTEX-14JIY-0011-R1a-SM-68713
60-69 years, male
45.5
GTEX-Q2AG-0011-R1A-SM-2HMJI
40-49 years, female
45.3
GTEX-13G51-0011-R1a-SM-5LZYR
60-69 years, male
45.0
GTEX-ZUA1-0011-R1a-SM-4YCE2
40-49 years, male
45.0
GTEX-RNOR-0011-R1A-SM-CNNP5
50-59 years, female
44.9
GTEX-YFC4-0011-R1a-SM-4V6EH
40-49 years, female
44.7
GTEX-14E7W-0011-R1b-SM-5YYAT
70-79 years, male
44.6
GTEX-131YS-0011-R1a-SM-5DUXC
60-69 years, female
44.4
GTEX-1IDJH-0011-R1b-SM-ARU6U
70-79 years, male
43.8
GTEX-PVOW-0011-R1A-SM-32PL6
40-49 years, male
43.4
GTEX-1EMGI-0011-R1b-SM-9OSWZ
60-69 years, male
43.4
GTEX-145MH-0011-R1b-SM-5PNWY
50-59 years, male
42.9
GTEX-R55E-0011-R1A-SM-2TC6N
20-29 years, male
42.3
GTEX-1IY9M-0011-R1a-SM-EXURZ
60-69 years, male
42.3
GTEX-12ZZX-0011-R1a-SM-5EQ6B
40-49 years, female
41.9
GTEX-1F6IF-0011-R1b-SM-9WG78
60-69 years, male
41.1
GTEX-13JUV-0011-R1a-SM-5LZY4
40-49 years, female
40.0
GTEX-T5JC-0011-R1A-SM-32PM6
20-29 years, male
38.8
GTEX-16YQH-0011-R1a-SM-7LT8R
30-39 years, male
38.5
GTEX-131XH-0011-R1a-SM-5DUWA
50-59 years, male
35.1
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HIPPOCAMPAL FORMATION - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.
Max subtype Scaled Tags Per Millioni
Max subtype Scaled Tags Per Million is the value for the subtype with the largest average Scaled Tags Per Million across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.
Normal distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. nTPM values of the individual samples are presented next to the box plot.
Max subtype nTPMi
Max subtype nTPM is the value for the subtype with the largest average nTPM across the subtype samples.